"Williams develops this new process philosophy of signs through a formal model, in contrast to earlier structuralist definitions. He draws on the philosophies of Deleuze and Whitehead, criticises earlier work on the sign in biology by Jakob von Uexküll, and connects to contemporary work on process in the philosophy of biology by John Dupré."
NDPR Review of Williams' A Process Philosophy of Signs HERE, also many good papers available on Deleuze by Williams at his website HERE.